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In 2015, a review recovered mDNA from ancient canid specimens which were discovered on Zhokhov Island along with the Yana river, arctic Siberia. These specimens included the mandible of a 360,000–400,000 YBP Canis cf. variabilis (wherever cf. is a Latin phrase indicating uncertain). Phylogenetic analyses of such canids uncovered 9 mDNA haplotypes